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Discovering Boulder County’s tiniest residents

Discovering Boulder County’s tiniest residents

CU Boulder alum and experienced caver Dave Steinmann recently discovered a new species of pseudoscorpion in Mallory Cave, with a moniker honoring its namesake hometown


When Dave Steinmann(Phys’90) first started classes at the University of Colorado Boulder in 1984, he had never explored a cave before and never really thought much about caves. However, when his new dorm-mate suggested they try his dad’s favorite hobby of caving, what seemed at first like an adventurous new pastime soon turned into a lifestyle for Steinmann—one that he has continued for more than 30 years and leading to his discovery of almost 100 new cave-dwelling species.

Steinmann, now a research associate with the Denver Museum of Nature & Science’s Zoology Department, most recently discovered a new species of pseudoscorpion named after the city closest to where it was found—none other than CU’s hometown of Boulder. Steinmann said that he knew almost immediately that the critter that is now known as Larca boulderica was a new species.

Nathan, Debbie and David Steinmann wearing caving helmets

Dave Steinmann (right) with his son, Nathan (left), and wife, Debbie (center), as they get ready to go caving. (Photo: Dave Steinmann)

When he first spotted it in Mallory Cave, one of Boulder’s most well-known cave systems thanks to its role in bat conservation, he immediately noticed its unique, almost lentil-shaped body and adaptations for cave living, such as its pale color. These specimens were later verified as a new species by Mark Harvey, a pseudoscorpion expert at the Western Australian Museum; Harvey and Steinmann recently in ZooKeys.

Steinmann notes that it’s typically not difficult to discern when a specimen is a new species, as it happens pretty frequently in the ancient cave systems right below our feet.

“I always say that if I want to discover a new species, I just need to visit a new cave,” he says.

Why are caves such a great place to make new discoveries? The answer lies in their role as a sort of refuge from climate change, Steinmann notes. In caves, insects can hide from the effects of temperature, floral and faunal changes that happen more rapidly in the outside world, facilitating isolated evolutionary changes.

Changing cave life

However, even cave life is changing. Lately, the temperature inside of caves, typically very cold, has been observed to be rising on a minuscule scale. Although this may seem trivial, even a few degrees’ difference can have immeasurable effects on the delicate life structures within the caves.

Similarly, outside temperatures affect which species go in and out of the cave systems, most notably bats. With the recent spike in white-nosed syndrome in bat populations, the number of bats in cave systems has decreased dramatically, with disastrous effects on internal cave species such as Larca boulderica, who survive on organic material—most often wood brought into the cave—and guano (bat fecal matter).

These changes are slow to progress, though, and there is still time to save cave ecosystems like that of Mallory Cave, which is closed to the public to protect the bat population inside (although it’s still possible to hike up to the cave entrance, a pleasant and short hike for anyone hoping to get outside).

So, how did Steinmann spot these teeny tiny bugs who live on bat feces? Well, after more than 30 years of experience, he has some tricks up his sleeve. One of the easiest methods he uses to spot tiny critters is simply by turning over rocks or pieces of wood.

When species like pseudoscorpions are disturbed by the movement or sense the carbon dioxide released by human breathing, they tend to skitter in every direction, looking for a new spot to curl up and revel in the damp darkness. When they move around, according to Steinmann, it’s just a game of whether you can catch them quickly enough.

a reddish-orange Larca boulderica pseudoscorpion

The newly described pseudoscorpion Larca boulderica is about the size of a sesame seed and is only known to live in Boulder, Colorado. (Photo: Dave Steinmann)

To catch samples, Steinmann usually brings simple tools along with him—a painter’s brush and some rubbing alcohol. When the brush is wetted with the alcohol, it’s easy to run it along a surface and pick up all of the tiny things residing there, including minuscule species of bugs likeLarca boulderica.

From there, it’s also easier to see what he’s found, as cave species are usually albino due to the lack of melanin— they don’t need pigmentation when there’s no sunlight—and they stand out against the dark ground and hairs of the paintbrush.

Looking for a gold bug

Despite being at it for multiple decades, Steinmann has no plans to slow down his caving career any time soon. He’s even made it a family pastime, and often spends time caving with his wife, Debbie, and his son, Nathan. He keeps an ongoing list of caves he plans to visit in the future and looks forward to making even more discoveries.

“I’d really like to find some kind of gold-colored bug and name it after the university,” he says, “or maybe even Coach Prime!”

He’s also enthusiastic about getting more students involved in caving, including caver and photographer Andres “Andy” Better, who will be a CU transfer student next fall. Steinmann emphasized how many different opportunities lie in the caving experience and says students of any background could find a niche interest in the hobby.

He also mentions local groups and clubs for both new and experienced cavers, including the Front Range Grotto and the Colorado Grotto, which meets at the Colorado School of Mines. He says that while anyone is welcome in caving, experienced members of the clubs can sometimes be protective of the places they visit, as human disturbances can harm delicate cave ecosystems, and caving as a hobby can be dangerous in a lot of ways.

However, if you’re looking to learn about caving with curiosity and respect, any of these clubs are great ways to get involved in this adventurous and exciting hobby—just be careful not to step in the bat guano because there could be a new species in there!


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